FAO - Key to subfamilies
of Gobiidae occurring in the Western Central
Pacific by Helen Larson
and E.O. Murdy
If you use this key please cite the following in publications:
Larson, H.K. and Murdy, E.O. 2001. Gobiidae. Gobies. Pp 3578-3603.
In: Carpenter, K.E. and Niem, V.H. (eds) FAO species
identification guide for fishery purposes.
The living marine resources of the western Central Pacific. Volume 6. Bony
fishes part 4 (Labridae to Latimeriidae). FAO, Rome.
1a. Dorsal and anal fins connected to caudal fin, both dorsal fins
united by
membrane; mud-burrowing, elongate gobies with pink to purple
skin.................................................................................................
Amblyopinae
1b. Dorsal and anal fins separated from caudal fin, both dorsal fins typically separate.................. 2
2a. Lower jaw typically possessing only a single row of teeth.................................................... 3
2b. Lower jaw typically possessing more than one row of teeth................................................. 4
3a. Pelvic frenum with fleshy lobes over spines; eyes lateral......................................... Sicydiinae
3b. Pelvic frenum without fleshy lobes; body elongate; eyes located mostly dorsally... Oxudercinae
4a. Paired anterior interorbital pores present or head pores completely
lacking. Pelvic frenum simple,
not folded forward, frenum without fleshy lobes around pelvic spines. If head
pores absent, then one or
more of the following conditions also exist: 1) pelvic frenum is present;
2) body fully scaled or mostly scaled;
and/or 3) no barbels present on chin [except for one genus, Gnatholepis,
only non-coral reef gobies are
included here. Gnatholepis possesses
head pores.].....................................
Gobionellinae (sensu Pezold, 1993)
4b. Usually a single anterior interorbital pore present or head pores
completely lacking. If two anterior
interorbital pores present, then pelvic frenum folded forward and a fleshy
lobe present around each
spine. If head pores absent, then one or more of the following conditions
also exist: 1) pelvic frenum is
absent; 2) body naked or with a few scales on caudal peduncle; and/or 3) barbels
present on chin
[although exceptions exist, head pores are typically absent only in a few
small coral reef
gobies]...............................................................................................
Gobiinae (sensu Pezold, 1993)
Key to the genera of Amblyopinae occurring in the area
(from Shibukawa and Murdy, unpubl.)
Note: There are
several known species that do not belong to any described genus; these are
not
included in the key (see species list).
1a. Pelvic fins moderate to large, pelvic fin length 64% or more of head
length (typically greater
than 70%); pelvic fins always fully united and forming a disc..........................
2 (Taenioides Group)
1b. Pelvic fins small to moderate, pelvic fin length 62% or less of head
length, (typically less
than 40%); pelvic fins fully united forming a disc (Caragobius
and Trypauchen), or emarginate
posteriorly (Amblyotrypauchen and Ctenotrypauchen),
or completely separate
(Trypauchenichthys).................................... 5 (Trypauchen
Group)
2a. Head and body with distinct cutaneous ridges; upper lip thick, fleshy,
with fringed dorsal margin;
caudal fin short, the length usually subequal to head length; ventral surface
of lower jaw usually with
several clusters of long, fleshy barbels; head and body entirely naked................................. Taenioides
2b. Head without distinct cutaneous ridges; upper lip thick or thin, not
fringed dorsally; caudal fin long or
very long, the length usually greater than head length (excluding Brachyamblyopus);
head with or without
barbels; minute cycloid scales at least on posterior part of body (sometimes
difficult to discern without
magnification).................... 3
3a. Most pectoral-fin rays simple, free from fin membrane; pectoral-fin rays 20-65, usually more than 23 4
3b. Most pectoral-fin rays branched, not forming free rays; pectoral-fin rays 23 or fewer Brachyamblyopus
4a. Head and body subcylindrical (slightly compressed posteriorly), greatly
elongate, body depth less than
10 % of standard length; teeth on outermost row of jaws enlarged, fang-like
(Fig. 5); a pair of symphysial canines
on lower jaw; scales minute, typically smaller than eye diameter (except for
those on posterior
half of body)..................................................................................................................
Odontamblyopus
4b. Head and body compressed, rather short, body depth 14.1-16.0 % of
standard length; teeth of
outermost row of jaws close-set, flattened, not fang-like; no symphysial canine
teeth on
lower jaw; scales large, distinctly larger than eye diameter...................................................
Pseudotrypauchen
5a. A pouch-like cavity at dorsal margin of operculum present (Fig. 4);
pectoral fin emarginate, asymmetrical
dorsoventrally, with large upper lobe; pore-like posterior nostril enlarged,
distinctly greater than eye; frontal crest
well-developed, prominent in external view, sometimes with serrated dorsal
margin (sometimes invisible in external
view); segmented caudal-fin rays 9+8, including 8+7 branched rays..............................................................
6
5b. No pouch-like cavity at dorsal margin of operculum; pectoral fin rounded,
symmetrical dorsoventrally;
pore-like posterior nostril subequal to eye diameter; dorsal margin of frontal
crest usually visible externally,
but not prominent; segmented caudal-fin rays usually 7+6, including 6+5 branched
rays....................... Caragobius
6a. Fang-like teeth on jaws; some scale patches on head.................. .Amblyotrypauchen
6b. No fang-like teeth on jaws (teeth on outer row sometimes strongly
caninoid); head typically naked ....7
7a. Pelvic fins separated to base; I,3 pelvic-fin rays; frontal crest
prominent with distinct serrated dorsal margin and
horn-like projection directed anteriorly (at least in adults).................................
Trypauchenichthys
7b. Pelvic fins united medially (sometimes emarginate posteriorly); I,5
pelvic-fin rays; frontal crest prominent, sometimes
with weak serration along dorsal margin, but not projecting anteriorly.....................................
8
8a. Belly fully scaled (sometimes with narrow naked area along ventral
midline), typically not possessing ventral
keel; pelvic fins united and rounded posteriorly, typically forming funnel-like
disc............. Trypauchen
8b. Belly naked, with prominent ventral keel; pelvic fins connected medially,
but
emarginate posteriorly, not forming a disc.............................................................................
Ctenotrypauchen
Key to the genera of Sicydiinae occurring in the area
1a. Comb-like or tricuspid teeth in either upper or lower jaw, conical teeth may be present also... 2
1b. No comb-like or tricuspid teeth in either jaw, only conical teeth present.................... Sicyopus
2a. Upper jaw with small tricuspid teeth at front and conical teeth at
side; lower jaw with
small horizontal teeth at front only and conical teeth behind..............................................
Lentipes
2b. Upper jaw with all tricuspid teeth; lower jaw with horizontal teeth
along whole length and conical
teeth behind .................................................................................................................3
3a. Gap in middle of upper jaw tooth rows; more than 50 scales in
lateral series......... Sicyopterus
3b. No gap in middle of upper jaw tooth rows which form continuous band
of tricuspid teeth; fewer than 50 scales in lateral series........................................................
Stiphodon
Key to the genera of Oxudercinae occurring in the area
1a. Lower eyelid (dermal cup) absent........................................................................... 2
1b. Lower eyelid (dermal cup) present......................................................................... 7
2a. Spinous dorsal fin with five spines........................................................................... 3
2b. Spinous dorsal fin with six spines............................................................................ 5
3a. Second dorsal fin with 23 or fewer total elements; anal fin with 23 or fewer total elements......
.................................................................................................................. Apocryptes
3b. Second dorsal fin with 27 or more total elements; anal fin with 26 or more total elements.........
.................................................................................................................................. 4
4a. Second dorsal fin 27-30; caudal-fin length typically more than 23% SL; head length greater than 22% SL .....Zappa
4b. Second dorsal fin I, 28-32; caudal-fin length 23% SL or less; head
length less than 22% SL........... Pseudapocryptes
5a. Second dorsal fin with 24 or fewer total elements, modally fewer;
anal fin with 23 or
fewer total elements; longitudinal scale count fewer than 60.............................................................
Apocryptodon
5b. Second dorsal fin with 24 or more total elements, modally more;
anal fin with 24 or more total
elements, modally more; longitudinal scale count typically more than 60...............................................................
6
6a. No prominent canine tooth lateral to upper jaw symphysis; head
length 24% SL or less; second
dorsal-fin base typically 45% SL or greater; caudal-fin length 19% SL or greater..........................
Parapocryptes
6b. Prominent canine tooth (about twice as long as others) on each
side of upper jaw symphysis;
head length 24% SL or greater; second dorsal-fin base 45% SL or less; caudal-fin
length 19% SL
or less..............................................................................................................................
Oxuderces
7a. Two canine teeth internal to lower jaw symphysis; anal-fin base and second dorsal-fin base 34% SL or greater 8
7b. No canine teeth internal to lower jaw symphysis; anal-fin base and second dorsal-fin base 27% SL or less 9
8a. Barbels present on underside of head................................................................... Scartelaos
8b. No barbels on underside of head................................................................. Boleophthalmus
9a. A single row of teeth in upper jaw; teeth blunt, not curved............................ Periophthalmus
9b. Two rows of teeth in upper jaw; outermost teeth large and curved............ Periophthalmodon
1a. Head pores absent............................................................................................................ 2
1b. Head pores present (sometimes only preopercular pores present)......................................... 8
2a. First element of second dorsal and anal fins is always unsegmented...................................... 3
2b. All elements of second dorsal and anal fins segmented........................................................ 6
3a. Papillae in interorbital space many, small, close-set, forming
long curved row around top
of each eye; mouth may be enlarged in males...........................................................................
4
3b. Papillae in interorbital space few, widely spaced in loose row
around each eye; mouth
extremely enlarged in males .............................................................................................
5
4a. Intestine long and coiled into three loops; pectoral rays 11-13;
restricted to
Australia
4b. Intestine simple, with one “S-bend” loop; pectoral rays 13-20; Indo-Pacific........ Mugilogobius
5a. 16 segmented caudal fin rays; several estuarine species, pale to
brownish with dark spots and
blotches .......................................................................................................Calamiana
(in part)
5b. 17 segmented caudal fin rays; one estuarine species, plain grey
colour..........................................................................................
Eugnathogobius
(in part)
6a. Body usually naked below first dorsal fin; head papillae in transverse
pattern; body
transparent or yellowish translucent in life (fresh and brackish waters)................................. Gobiopterus
6b. Body usually fully scaled; head papillae usually in longitudinal pattern; body with dusky bands or spots 7
7a. Adults small, up to 25 mm SL; robust-bodied gobies with distinctive
banded color pattern
(brackish to freshwater)................................................................................................
Brachygobius
7b. Adults very small, at most reaching 15 mm SL; slender-bodied gobies
with one dusky band extending
from first dorsal fin, remainder of body with incomplete bands and blotches
(brackish water)................. Pandaka
8a. None or one pair of pores present on snout......................................................................... 9
8b. Two pairs of pores present on snout................................................................................. 16
9a. Body naked, mostly freshwater.................................................................. Schismatogobius
9b. Body scaled, at least on its posterior half; brackish to freshwater........................................ 10
10a. 17 segmented caudal fin rays.......................................................................................... 12
10b. 16 segmented caudal fin rays.......................................................................................... 11
11a. Mouth terminal, enlarged in males; gut simple, forming “S-bend”
loop; headpores usually
absent .....................................................................................................Calamiana
(in part)
11b. Mouth small, usually subterminal, with rounded snout overhanging
mouth; some headpores always
present; entire gut spirally coiled about its longitudinal axis....................................................
Pseudogobius
12a. Head papillae longitudinal; if transverse rows present, then lateral
canal over preopercle
and/or opercle present also; coloring variable..............................................................................................
13
12b. Head papillae with many transverse rows; head pores typically
present but lateral
canal over opercle always absent; distinctively spotted species............................................................
Stigmatogobius
13a. Preopercular pores and lateral canal present or absent (depending
on species); gut short, with
only 2-3 loops; jaws may be greatly enlarged in males, lips not reduced and
thin....................................................
14
13b. Preopercular pores and lateral canal always absent; gut long and
coiled into many loops
(12 or more); mouth small, lower lip reduced, thin and folded forward.................................................
Hemigobius
14a. No pores over top of opercle (one present above rear margin of
preopercle); preopercular pores
present or absent; second dorsal and anal fins with equal numbers of rays
or soft dorsal with one more
ray than anal...........................................................................................................................
15
14b. Two pores present over top of opercle (as well as one present
above rear margin of preopercle;
three preopercular pores usually present; anal fin with one or two more rays
than in second dorsal fin............. Rhinogobius
15a. Head depressed; body cylindrical or somewhat elongate; mouth large,
especially in
males, reaching beyond middle of eye................................................................................
Eugnathogobius
(in part)
15b. Head compressed, body often compressed, few species elongate;
mouth small and
terminal in females, large and inferior to subinferior in males..................................................................
Redigobius
16a. Shoulder girdle under gill cover with distinct fleshy lobes (1-4 finger-like flaps).................. 17
16b. Shoulder girdle under gill cover smooth or with minute bumps........................................... 18
17a. Head broader than deep; mouth inferior with fleshy lips; predorsal
scale count
16-42; body with rows of blotches and spots..........................................................................................
Awaous
17b. Head compressed, narrower than deep; mouth terminal, lips not
particularly fleshy; predorsal scale
count 0-23; body with variably developed transverse bands........................................................................
Stenogobius
18a. Cheek with large scales; teeth at sides of upper jaw directed
medially; anterior interorbital pore
paired; mouth horizontal, inferior (coral reefs)...............................................................................................
Gnatholepis
18b. Cheek naked; teeth at sides of jaws vertical or directed posteriorly................................... 19
19a. Median membranous crest or ridge usually present on nape; teeth
in upper jaw usually in a single row, may have
a few teeth on inner row anteriorly; eye with or without fleshy knob or tentacle;
tongue rounded .......Oxyurichthys
19b. No crest or ridge on nape; typically, teeth in upper jaw in two
or three rows; eye always
without fleshy knob or tentacle; tongue truncate.....................................................................................
Oligolepis
Key to the described genera of Gobiinae occurring in the area
NOTE: Many genera remain to be adequately
revised or defined; consequently the key is not guaranteed
to work for all species of a genus. Some genera will key out in more than
one place in the key. There are
several known species which do not belong to any described genus; these are
not included in the key.
1a. First gill slit closed by membrane; distinctive transverse papillae pattern on head (coral reefs) Hetereleotris
1b. First gill slit open; papillae pattern longitudinal or transverse.................................................. 2
2a. Body naked or with a few scales on caudal peduncle........................................................... 3
2b. Body scaled at least on posterior half.................................................................................. 8
3a. Pelvic fins separate and slender......................................................................................... 4
3b. Pelvic fins united............................................................................................................... 5
4a. Body deep, robust, eyes small (coral reefs)...................................................... Austrolethops
4b. Body slender, eyes moderate to large (coral reefs)................................. Trimmatom (in part)
5a. Teeth tricuspid; body slender (rocky shores)....................................................... Kelloggella
5b. Teeth pointed.................................................................................................................... 6
6a. Body short and/or compressed; anal fin with 10 or fewer elements, including spine................ 7
6b. Body elongate; anal fin with 13 elements, including spine (sandy shores).......... Parkraemeria
7a. Head and body deep and compressed; body and fins with thick mucous
coat; pelvic
fins short and fleshy (coral reefs)..............................................................................................
Gobiodon
7b. Body short and robust, compressed posteriorly but head usually
rounded; mucous coat
not greatly developed; pelvic fins not fleshy (deepwater)........................................................
Lubricogobius
8a. Thin dermal crest on top of head anterior to first dorsal fin................................................... 9
8b. No dermal crest anterior to first dorsal fin......................................................................... 11
9a. Sensory papillae on head transverse; body relatively plain dark
brown, with dark
blotch on shoulder just above pectoral fin base (estuaries)....................................................
Lophogobius
(in part)
9b. Sensory papillae on head longitudinal; colour pattern variable, often with small dark spots..... 10
10a. Dermal crest low, less than pupil diameter; body elongate; soft
dorsal and anal
rays I, 12 (estuaries, shallow reefs)...................................................................................
Cryptocentroides
10b. Dermal crest high, more than pupil diameter; body deep; soft dorsal and anal rays I,9 (estuaries) .......Cristatogobius
11a. Barbels present on ventral surface of head (may be on chin only),
barbels distinctly larger than
any elongate papillae..............................................................................................................12
11b. Papillae on underside of head may be elongate, but no barbels present.............................. 16
12a. Large black spot present dorsally on caudal fin; cheek and opercle
covered with scales
(deepwater) ..............................................................................................
Parachaeturichthys
12b. No large black spot on caudal fin; no scales on cheek or opercle....................................... 13
13a. One or two distinct folds on cheek; headpores present or absent;
barbels in several pairs or
small groups ............................................................................................................................15
13b. No folds on cheek; headpores always present, including one or
two over opercle; barbels profuse
or one pair only .........................................................................................................................14
14a. Barbels on head profuse, slender, forming fringe around head (coral reefs)..... Barbuligobius
14b. Barbels reduced to small pair on chin (freshwater to estuaries)........... Glossogobius (in part)
15a. Barbels only present on chin; entire fish greatly dorsoventrally
flattened and elongate
(deep reefs) ..................................................................................................Platygobiopsis
15b. Barbels may be present on chin, snout and sides of head; fish
stout, with depressed head
but body never extraordinarily elongate (coral reefs, estuaries)......................................... Gobiopsis
16a. First spine in first and second dorsal fins rigid and pungent, and usually thickened............... 17
16b. Dorsal-fin spines thin and flexible, not pungent or thickened.............................................. 21
17a. Preopercle with 1-3 flat spines; nape may have small scales (coral reefs)........... Oplopomus
17b. Preopercle without spines; nape scales may be as large as body scales (or absent)............ 18
18a. Nape scaled, at least opercle partly scaled, preopercle may or
may not be scaled; first spines of
first and second dorsal fins twice the thickness of remaining fin spines;
body scales ctenoid.......... 19
18b. Nape, preopercle and opercle naked; first spine of first and second
dorsal fins not much
" thicker than remaining spines; all scales cycloid (coral reefs).....................................
Echinogobius
19a. Preopercle and opercle fully scaled (deepwater)...................................................... Hazeus
19b. Preopercle naked, opercle partly scaled........................................................................... 20
20a. Opercle naked below level of upper pectoral-fin base (sand, coral reefs)........... Oplopomops
20b. Opercle scaled at least to level of upper one-third of pectoral-fin base (shallow reefs).. Opua
21a. Preopercle with a single large prominent spine (shallow reefs)......................... Gladiogobius
21b. Preopercle with 0-9 small spines..................................................................................... 22
22a. Preopercle with 1-9 spines (coral reefs)........................................................ Asterropteryx
22b. No spines on preopercle................................................................................................. 23
23a. First dorsal fin elongate, fin origin at rear of head opposite
rear end of opercle
(coral reefs) .......................................................................................................Discordipinna
23b. First dorsal fin shape variable, fin origin behind pectoral base............................................ 24
24a. Cheeks with papillae in transverse pattern, papillae prominent,
raised upon fleshy flaps (coral reefs,
estuaries)........................................................................................................................ Callogobius
24b. Cheeks with papillae in transverse or longitudinal pattern but
without prominent vertical fleshy flaps
bearing papillae..........................................................................................................................25
25a. Pelvic fins with thickened lobe around each pelvic spine, frenum folded forward forming a pocket....... 26
25b. Pelvic fins with or without thickened skin around each pelvic
spine, frenum flat, may be fleshy but
not folded forward ................................................................................................................
30
26a. Interobital canals separate, two anterior interorbital pores present..................................... 28
26b. Interorbital canal single, one (rarely two) anterior interobital pore present.......................... 27
27a. Edge of lower lip fused to underside of head, lip free at chin only (coral reefs)...... Luposicya
27b. Edge of lower lip free at sides, fused at chin (coral reefs).................................. Pleurosicya
28a. Pectoral rays all branched; eyes small (deepwater)......................................... Lobulogobius
28b. Pectoral fins with lower 2-6 rays unbranched and tips usually thickened; eyes large........... 29
29a. Gill opening wide and not attached to isthmus; head broad and
flattened; nape scaled
(coral reefs) .........................................................................................................Phyllogobius
29b. Gill opening wide or narrow, but always attached to isthmus; head
not flattened but
may be elongate; nape usually naked (coral reefs)...................................................
Bryaninops
30a. Chin with curved mental frenum (free fleshy flap) or distinct rounded to triangular knob..... 31
30b. Chin relatively smooth, without a mental frenum, may be slight
swelling on chin just anterior
to row of sensory papillae......................................................................................................
34
31a. Tips of upper pectoral-fin rays free and silk-like and no curved
canine tooth in
each side of lower jaw (usually shallow reefs).........................................................
Bathygobius
31b. Tips of upper pectoral-fin rays not free or if free, then a curved
canine tooth present at each
side of lower jaw ..............................................................................................................32
32a. Mouth subterminal with snout partly overhanging upper lip; soft
dorsal I,10-11, anal I,9-10 (coral reefs, estuaries)............................................
Istigobius (in part)
32b. Mouth terminal; soft dorsal I,7-9, anal I,6-9...................................................................... 33
33a. Head depressed; tongue bilobed or deeply concave; pelvic frenum
reduced or absent;
colouring usually white to yellowish with few dark markings (coral reefs,
sand)................ Cabillus
33b. Head not depressed; tongue blunt to rounded; pelvic frenum always
present, conspicuous;
colouring usually mottled, spotted and barred with brown (coral to rocky reefs)................... Palutrus
34a. Head rounded, the ventral surface scattered with numerous small
bumps, sides and top of
head covered with fleshy bumps or fine flaps (which may be close together);
nape naked
(coral reefs)............................................................................................................
Paragobiodon
34b. Head without fine fleshy flaps and bumps, nape scaled or naked....................................... 35
35a. Head pores absent; size not greater than 50 mm SL......................................................... 36
35b. Head pores present........................................................................................................ 39
36a. Head papillae conspicuous, in rows which may form ridges; head depressed (coral reefs)......
........................................................................................................................................ Feia
36b. Papillae on head small, not forming ridges; head compressed or cylindrical........................ 37
37a. Pelvic fin rays all unbranched, or at least fifth ray
unbranched (coral reefs)..........................
................................................................................................................ Trimmatom (in part)
37b. Pelvic fin rays branched; body scaled.............................................................................. 38
38a. Gill opening ends below rear margin of preopercle; head usually
broader than deep; vertical
bars with dark borders present on head at least (coral reefs, deepwater)................................ Priolepis
38b. Gill opening extends to below eye or at least to preopercular
margin; head usually deeper
than broad; colour pattern variable, often with spots on head (coral reefs).................................. Trimma
39a. Pelvic fins completely separate, no membrane connecting bases of fifth pelvic rays........... 40
39b. Pelvic fins partly or completely connected by membrane.................................................. 43
40a. Papillae on cheek include short transverse rows; gill opening
extends
to below preopercle (coral reefs)....................................................................
Amblyeleotris (in part)
40b. Papillae on cheek in longitudinal pattern........................................................................... 41
41a. Teeth in upper jaw in single row; adult size greater than 50 mm SL (coral reefs)
.......................................................................................................................... Valenciennea
41b. Teeth in upper jaw in two or more rows; adult size less than 30 mm SL............................. 42
42a. Fifth pelvic ray unbranched, usually considerably reduced, fin
rays
many-branched, often fringe-like (coral reefs)..................................................................
Eviota
42b. Fifth pelvic fin ray branched, fin rays branched at tips, but
not fringe-like (coral reefs,
deepwater) ......................................................................................................Sueviota
(in part)
43a. If any papillae rows on head are on raised fleshy ridges, then
mouth not small and nearly
vertical ................................................................................................................................44
43b. Usually at least two rows of papillae on head on raised, fleshy,
longitudinal
ridges; mouth small, oriented nearly vertically (estuaries).......................................... Mangarinus
44a. Cheeks and operculum covered with scales (may be small or embedded).......................... 45
44b. Cheeks and operculum partially scaled or naked (may be small or embedded).................... 48
(deepwater).....................................................................................
Egglestonichthys
(in part)
45b. Headpores present; frenum between pelvic spines present.................................... 46
46a. Gill opening restricted to pectoral base or to below opercle;
if transverse papillae present
under eye, then snout rounded and may overhang upper lip slightly.................................................
47
46b. Gill opening very wide, extending up to below eye; 7-10 short
rows of transverse papillae below eye;
snout pointed, with lower jaw tip anteriormost; 5-6 brown spots along side
of body (estuaries, coastal).... Isthmogobius
47a. Body slender, body depth contained more than four times in SL;
prominent recurved
canine tooth at angle of lower jaw (coral reefs).......................................................
…..Macrodontogobius
47b. Body deep, body depth contained less than four times in SL; no
prominent recurved
canine tooth on lower jaw (coral reefs, estuaries) ........................................... …..…………….Exyrias
48a. Gill opening extending to below rear margin of preopercle (or farther forward)...... 49
48b. Gill opening restricted to pectoral-fin base or slightly further forward to below opercle........... 63
49a. Head papillae in transverse pattern, at least transverse rows present under eye...... 55
49b. Head papillae in longitudinal pattern, some rows may be very short....................... 50
50a. Anal fin always with one or more segmented ray than in soft dorsal
(sandy shores,
estuaries) .......................................................................................................Silhouettea
50b.
Anal and soft dorsal fins with equal numbers of rays, or dorsal fin with one
or more ray
than anal fin .................................................................................................................
51
51a.
Iris lappet present in eye; tongue deeply bilobed; gill opening wide (nearly
to eye) and free
of isthmus (coral reefs, estuaries).........................................................................
Psammogobius
51b.
No iris lappet present; tongue may be concave but not deeply bilobed; gill
opening
variable.........................................................................................................................52
52a.
Soft dorsal and anal rays I,8-10, most papillae rows on cheek long, not reduced
(estuaries, freshwater)...............................................................................
Glossogobius
(in part)
52b. Soft dorsal and anal rays I,10-12; some cheek papillae rows short or broken.......... 53
53a. A distinct black ocellus in each dorsal fin (coral reefs)........................... Signigobius
53b. No distinct black ocellus in each dorsal fin........................................................... 54
54a.
Caudal fin always rounded, shorter than head; body pale with dark spots and
bright
white spot on pectoral fin (coral reefs)......................................................................
Ctenogobiops
54b.
Caudal fin usually pointed, longer than head; body with bars, spots and/or
stripes,
no bright white spot on pectoral fin (coral reefs)........................................................
Vanderhorstia
55a. Scales small; 44 or more in a lateral series........................................................... 57
55b. Scales large; 40 or fewer in a lateral series.......................................................... 56
56a.
First dorsal fin tall and broad with dark bands and/or spots; body with dark
oblique bars
(estuaries, coral reefs)................................................................................
Mahidolia
56b.
First dorsal rounded to triangular, relatively unpatterned; body with spots
and blotches
but not dark oblique bands (estuaries, mangroves).................................... Acentrogobius
(in part)
57a. Teeth present on vomer (which is curved into mouth); body pale with distinct dark markings (coral reefs)............................................................................................................... Stonogobiops
57b. No teeth on vomer (note: vomer itself may be curved into mouth); colour variable................. 58
58a. Second dorsal fin rays I,12-14; anal fin rays I,12-14 (coral reefs)......................... Amblyeleotris (in part)
58b. Second dorsal fin rays I,9-11; anal fin rays I,8-10................................................. 59
59a.
Preopercular pores absent; vertical white lines over abdomen present, especially
prominent
in males (shallow reefs)........................................................................
Psilogobius
(in part)
59b. Preopercular pores present; no vertical white abdominal lines present.................... 60
60a. No posterior oculoscapular canal or pores over top of opercle............................... 61
60b. Posterior oculoscapular canal and two pores present over top of opercle............... 62
61a.
First dorsal fin broad and\or tall, longer than or equal to body depth; if
body dark, no bright
white stripe along nape midline (shallow reefs, estuaries)....................................... Myersina
(in part)
61b.
First dorsal fin low, with black ocellus; body dark with nape and top of head
bright white
(coral reefs).................................................................................................................
Lotilia
62a.
Gill opening wide, membranes forming distinct free fold across isthmus, attaching
to isthmus in
front of fold (shallow reefs, estuaries)...................................................................
Myersina
(in part)
62b.
Gill opening moderately wide, membranes not forming fold across isthmus, but
attaching
to sides of isthmus (coral reefs, deepwater)......................................................... Cryptocentrus
63a. Papillae on cheek in transverse pattern............................................................................ 64
63b. Papillae on cheek in longitudinal pattern........................................................................... 68
64a. Scales present on cheek, at least behind eye.................................................................... 65
64b. Cheek naked................................................................................................................. 66
65a. Upper half of cheek covered with scales, nape scales extend forward
to snout
(deepwater)...........................................................................................
Egglestonichthys (in
part)
65b. Scales on cheek restricted to patch behind eye, nape scales
extend up to behind eyes (reefs, estuaries, deepwater)................................................
Acentrogobius (in part)
66a. Large posterolaterally-directed canine tooth on lower jaw, often
visible when mouth closed;
snout rounded and overhangs upper lip in most species (coral reefs)................................
Amblygobius
66b. No large posterolaterally-directed canine tooth on lower jaw, snout not overhanging upper lip 67
67a. Nape naked; vertical papillae rows on mid-cheek do not extend
ventrally past lowermost longitudinal cheek row (estuaries)..........................................
Lophogobius (in part)
67b. Nape usually with scales, midline may be naked; one vertical papilla
row on mid-cheek extends
ventrally past lowermost longitudinal cheek row (shallow reefs, estuaries).............................. Drombus
68a. Cheek (and opercle) with scales; caudal fin long and slender (deepwater)...... Obliquogobius
68b. Cheek naked, opercle scaled or naked............................................................................. 69
69a.
Cheeks swollen; mouth large, extends at least to rear edge of eye (may extend
well past eye);
gill opening restricted to pectoral base (coral reefs)................................ Tomiyamichthys
69b. Cheeks not swollen; mouth reaches to below eye; gill opening to under opercle or to rear of eye..... 70
70a. No oculoscapular canal over opercle, preopercle with only two
or no pores (coral reefs,
deepwater) ....................................................................................................Sueviota
(in part)
70b. Oculoscapular canal present over opercle and preopercle with three pores........................ 71
71a. Pore behind eye long and slit-like (estuaries, deepwater).................................... Aulopareia
71b. Pore behind eye rounded................................................................................................ 72
72a.Mouth terminal; snout pointed or rounded, tip not overhanging upper
lip; no free flap or
frenum on lower jaw, skin joining lower lip to isthmus..............................................................
73
72b. Mouth subterminal; snout rounded to blunt, with tip slightly overhanging
upper lip; free flap or
frenum on lower jaw in front of isthmus may be visible (coral reefs, estuaries).... Istigobius
(in part)
73a. Pre-pelvic area naked; snout relatively long, depressed and pointed;
jaws ending below rear half of
eye; cheek and opercle naked; 0-4 predorsal scales present (may be embedded);
second dorsal
rays I,8 (estuarine)..................................................................................................Afurcagobius
73b. Pre-pelvic area with some scales, may be large and deciduous;
snout short, rounded to pointed, not
depressed; jaws may end below anterior part of eye or mid-eye; cheek and opercle
naked or
partly scaled; predorsal scales present or absent; second dorsal rays I,7-11......................... 74
74a. Sensory papillae rows on cheek very short, papillae small and
few; head pointed in dorsal view;
eyes large and set high on side of head; pelvic fins may have frenum reduced
or absent; membrane
between fifth pelvic rays may be reduced, pelvic fins separate in some species;
first dorsal fin
generally triangular and conspicuously marked; when live, body translucent
with black, brown and white markings (coral reefs).................................................
Fusigobius
74b. Most sensory papillae rows on cheek long (or multiple) and conspicuous;
head may be pointed or
rounded in dorsal view; eye variable in size but usually not large and set
high on side of head; pelvic
fins always with frenum present; fifth pelvic rays always joined by membrane;
first dorsal fin shape
variable, but generally without conspicuous black marks; when live, body opaque,
colour variable..........................................................................................................
75
75a. Single row of papillae usually present across rear of chin, if
large patch of papillae present on
chin, then some papillae rows on side of head multiple, row directly under
eye always single; snout
usually rounded; scales may be present on opercle or on cheek behind eye;
dorsal spines may be
long and at least second spine filamentous in both sexes; body colour pattern
variable (reefs,
estuaries, deepwater)......................................................
Acentrogobius
(in part)
75a. Large patch of papillae on chin; on cheek, multiple short irregular
rows of papillae (may be
vertically oriented) present between two lowermost longitudinal papillae rows
and papillae row
directly under eye double, OR 5-6 longitudinal rows of papillae on cheek,
none multiple; first
or second dorsal spine may be elongate in mature males; snout short and pointed;
no scales on
cheek or opercle; body whitish to pale yellowish with fine dark spots and
speckles, dark spots
along mid-side of body smaller than eye (sandy habitats near shallow reefs
and estuaries)............. Favonigobius
List of marine and brackish
water species occurring in the area.
Species which live in freshwater but may occur in estuaries as adults or larvae are marked by
an asterisk (*). Listed are 105 genera (including three new genera for described species of
amblyopines), and 534 species (undescribed species not included).
Amblyotrypauchen arctocephalus (Alcock, 1890)
Brachyamblyopus brachysoma (Bleeker, 1853)
Caragobius coecus (Weber, 1913)
Caragobius geomys Fowler, 1935.
Caragobius monserrati (Roxas and Ablan, 1940)
Caragobius olivaceus (Herre, 1927)
Caragobius rubristriatus (Saville-Kent, 1889)
Caragobius typhlops Smith and Seale, 1906
Caragobius urolepis (Bleeker, 1852)
Ctenotrypauchen microcephalus (Bleeker, 1860)
New genus (A) jacksoni (Smith, 1943)
New genus (B) nigrimarginatus (Hora, 1924)
New genus (C) sumatranus (Volz, 1903)
Odontamblyopus rubicundus (Hamilton-Buchanan,
1822)
Odontamblyopus tenuis (Day,
1876)
Pseudotrypauchen multiradiatus Hardenberg, 1931
Taenioides anguillaris (Linnaeus, 1758)
Taenioides caniscapulus Roxas and Ablan, 1938
Taenioides cirratus (Blyth, 1860)
Taenioides eruptionis (Bleeker,
1849)
Taenioides gracilis (Valenciennes, 1837)
Taenioides mordax (De Vis,
1883)
Taenioides purpurascens (De Vis, 1884)
Trypauchen totoyensis (Garman, 1903)
Trypauchen raha Popta, 1922
Trypauchen taenia Koumans, 1953
Trypauchen vagina (Bloch and Schneider, 1801)
Trypauchenichthys sumatrensis Hardenberg, 1931
Trypauchenichthys typus (Bleeker, 1860)
Subfamily GOBIINAE
Acentrogobius audax Smith, 1959
Acentrogobius bifrenatus (Kner, 1856)
Acentrogobius caninus (Valenciennes, 1837)
Acentrogobius frenatus (Günther, 1861)
Acentrogobius gracilis (Bleeker, 1875)
Acentrogobius janthinopterus (Bleeker, 1852)
Acentrogobius leftwichi (Ogilby, 1910)
Acentrogobius madraspatensis (Day, 1868)
Acentrogobius moloanus (Herre, 1927)
Acentrogobius nebulosus (Forsskal, 1775)
Acentrogobius pyrops (Whitley, 1954)
Acentrogobius suluensis (Herre, 1927)
Acentrogobius viganensis (Steindachner, 1893)
Acentrogobius viridipunctatus (Valenciennes, 1837)
Afurcagobius tamarensis (Johnston, 1883)
Amblyeleotris aurora (Polunin and Lubbock, 1977)
Amblyeleotris callopareia Polunin and Lubbock, 1979
Amblyeleotris delicatulus Smith, 1958
Amblyeleotris diagonalis Polunin and Lubbock, 1979
Amblyeleotris exilis (Smith, 1958)
Amblyeleotris fasciata (Herre, 1953)
Amblyeleotris fontanesii (Bleeker, 1852)
Amblyeleotris guttata (Fowler, 1938)
Amblyeleotris gymnocephala (Bleeker, 1853)
Amblyeleotris japonica Takagi, 1957
Amblyeleotris latifasciata Polunin and Lubbock, 1979
Amblyeleotris macronema Polunin and Lubbock, 1979
Amblyeleotris novaecaledoniae Goren, 1981
Amblyeleotris ogasawarensis Yanagisawa, 1978
Amblyeleotris periophthalma (Bleeker, 1853)
Amblyeleotris randalli Hoese and Steene, 1978
Amblyeleotris rhyax Polunin and Lubbock, 1979
Amblyeleotris steinitzi (Klausewitz, 1974)
Amblyeleotris sungami (Klausewitz, 1969)
Amblyeleotris wheeleri Polunin and Lubbock, 1977
Amblyeleotris yanoi Aonuma and Yoshino, 1996
Amblygobius buanensis (Herre, 1927)
Amblygobius bynoensis (Richardson, 1844)
Amblygobius decussatus (Bleeker, 1855)
Amblygobius esakiae (Herre, 1939)
Amblygobius hectori (Smith, 1956)
Amblygobius linki Herre, 1927
Amblygobius nocturnus (Herre, 1945)
Amblygobius phalaena (Valenciennes, 1837)
Amblygobius rainfordi (Whitley, 1940)
Amblygobius sphynx (Valenciennes, 1837)
Asterropteryx bipunctatus Allen and Munday, 1995
Asterropteryx ensiferus (Bleeker, 1874)
Asterropteryx semipunctatus (Rüppell, 1830)
Asterropteryx spinosus (Goren, 1981)
Asterropteryx striatus Allen and Munday, 1995
Aulopareia atripinnatus (Smith, 1931)
Aulopareia cyanomos (Bleeker, 1849)
Aulopareia janetae Smith, 1945
Aulopareia koumansi (Herre, 1937)
Aulopareia spilopterus (Smith, 1932)
Aulopareia unicolor (Valenciennes, 1837)
Austrolethops wardi Whitley, 1935
Barbuligobius boehlkei Lachner and McKinney, 1974
Bathygobius albopunctatus (Valenciennes, 1837)
Bathygobius coalitus (Bennett, 1832)
Bathygobius cocosensis (Bleeker, 1854)
Bathygobius cotticeps (Steindachner, 1880)
Bathygobius cyclopterus (Valenciennes, 1837)
Bathygobius fuscus (Rüppell, 1830)
Bathygobius kreffti (Steindachner, 1866)
Bathygobius laddi (Fowler, 1931)
Bathygobius meggitti (Hora and Mukerji, 1936)
Bathygobius padangensis (Bleeker, 1851)
Bathygobius panayensis (Jordan and Seale, 1907)
Bathygobius petrophilus (Bleeker, 1853)
Bryaninops amplus Larson, 1985
Bryaninops diannae Larson, 1985
Bryaninops erythrops (Jordan and Seale, 1906)
Bryaninops isis Larson, 1985
Bryaninops loki Larson, 1985
Bryaninops natans Larson, 1985
Bryaninops nexus Larson, 1987
Bryaninops ridens Smith, 1959
Bryaninops tigris Larson, 1985
Bryaninops yongei (Davis and Cohen, 1969)
Cabillus lacertops Smith, 1959
Cabillus macrophthalmus (Weber, 1909)
Cabillus tongarevae (Fowler, 1927)
Callogobius bauchotae Goren, 1979
Callogobius centrolepis Weber, 1909
Callogobius clitellus McKinney and Lachner, 1978
Callogobius crassus McKinney and Lachner, 1984
Callogobius depressus (Ramsay and Ogilby, 1886)
Callogobius flavobrunneus (Smith, 1958)
Callogobius hasselti (Bleeker, 1851)
Callogobius hastatus McKinney and Lachner, 1978
Callogobius liolepis Koumans, 1931
Callogobius maculipinnis (Fowler, 1918)
Callogobius okinawae Snyder, 1908)
Callogobius sclateri (Steindachner, 1880)
Callogobius stellatus McKinney and Lachner, 1978
Callogobius tanegasimae (Snyder, 1908)
Cristatogobius albius Tchaw-ren, 1959
Cristatogobius lophius Herre, 1927
Cristatogobius nonatoae (Ablan, 1940)
Cryptocentroides cristatus (Macleay, 1881)
Cryptocentroides insignis (Seale, 1910)
Cryptocentrus albidorsus Yanagisawa, 1978
Cryptocentrus bulbiceps (Whitley, 1953)
Cryptocentrus caeruleomaculatus (Herre, 1933)
Cryptocentrus cebuanus Herre, 1927
Cryptocentrus cinctus (Herre, 1936)
Cryptocentrus cyanotaenia (Bleeker, 1853)
Cryptocentrus diproctotaenia (Bleeker, 1876)
Cryptocentrus fasciatus (Playfair, 1866)
Cryptocentrus filifer (Valenciennes, 1837)
Cryptocentrus inexplicatus (Herre, 1934)
Cryptocentrus insignitus (Whitley, 1956)
Cryptocentrus leonis Smith, 1931
Cryptocentrus leptocephalus Bleeker, 1876
Cryptocentrus leucostictus (Gunther, 1872)
Cryptocentrus lutheri Klausewitz, 1960
Cryptocentrus maudae Fowler, 1937
Cryptocentrus niveatus (Valenciennes, 1837)
Cryptocentrus obliquus (Herre, 1934)
Cryptocentrus pavoninoides (Bleeker, 1854)
Cryptocentrus pretiosus (Rendahl, 1924)
Cryptocentrus shigensis Kuroda, 1956
Cryptocentrus strigilliceps (Jordan and Seale, 1906)
Cryptocentrus wehrlei Fowler, 1937
Ctenogobiops aurocingulus (Herre, 1935)
Ctenogobiops crocineus Smith, 1959
Ctenogobiops feroculus Lubbock and Polunin, 1977
Ctenogobiops pomastictus Lubbock and Polunin, 1977
Ctenogobiops tangaroai Lubbock and Polunin, 1977
Discordipinna griessingeri Hoese and Fourmanoir, 1978
Drombus dentifer Hora, 1923
Drombus globiceps (Hora 1923)
Drombus halei (Whitley, 1935)
Drombus kranjiensis (Herre, 1940)
Drombus ocyurus Jordan and Seale, 1906
Drombus simulus (Smith, 1960)
Drombus triangularis (Weber, 1911)
Echinogobius hayashii Iwata, Hosoya and Niimura, 1998
Egglestonichthys bombylios Larson and Hoese, 1997
Egglestonichthys melanoptera (Rao, 1971)
Eviota afelei Jordan and Seale, 1906
Eviota albolineata Jewett and Lachner, 1983
Eviota bifasciata Lachner and Karnella, 1980
Eviota cometa Jewett and Lachner, 1983
Eviota distigma Jordan and Seale, 1906
Eviota fasciola Karnella and Lachner, 1981
Eviota herrei Jordan and Seale, 1906
Eviota infulata (Smith, 1956)
Eviota irrasa Karnella and Lachner, 1981
Eviota lachdeberei Giltay, 1933
Eviota latifasciata Jewett and Lachner, 1983
Eviota melasma Lachner and Karnella, 1980
Eviota monostigma Fourmanoir, 1971
Eviota nebulosa Smith, 1958
Eviota nigriventris Giltay, 1933
Eviota pellucida Larson, 1976
Eviota prasina (Klunzinger, 1871)
Eviota prasites Jordan and Seale, 1906
Eviota pseudostigma Lachner and Karnella, 1980
Eviota punctulata Jewett and Lachner, 1983
Eviota queenslandica Whitley, 1932
Eviota saipanensis Fowler, 1945
Eviota sebreei Jordan and Seale, 1906
Eviota sigillata Jewett and Lachner, 1983
Eviota smaragdus Jordan and Seale, 1906
Eviota sparsa Jewett and Lachner, 1983
Eviota spilota Lachner and Karnella, 1980
Eviota storthynx (Rofen, 1959)
Eviota variola Lachner and Karnella, 1980
Eviota zebrina Lachner and Karnella, 1978
Eviota zonura Jordan and Seale, 1906
Exyrias belissimus (Smith, 1959)
Exyrias ferrarisi Murdy, 1985
Exyrias puntang (Bleeker, 1851)
Favonigobius exquisitus Whitley, 1950
Favonigobius lentiginosus (Richardson, 1844)
Favonigobius melanobranchus (Fowler, 1934)
Favonigobius opalescens (Herre, 1936)
Favonigobius reichei (Bleeker, 1853)
Feia nympha Smith, 1959
Fusigobius duospilos Hoese and Reader, 1985
Fusigobius longispinus Goren, 1978
Fusigobius neophytus (Günther, 1877)
Fusigobius signipinnis Hoese and Obika, 1988
Gladiogobius ensifer Herre, 1933
Glossogobius aureus Akihito and Meguro, 1975
Glossogobius bicirrhosus (Weber, 1894)
Glossogobius celebius (Valenciennes, 1837)
Glossogobius circumspectus (Macleay, 1883)
Glossogobius giuris (Hamilton, 1822)
Glossogobius sparsipapillus Akihito and Meguro, 1976
Gobiodon acicularis Harold and Winterbottom, 1995
Gobiodon albofasciatus Sawada and Arai, 1972
Gobiodon atrangulatus Garman, 1903
Gobiodon axillaris De Vis, 1884
Gobiodon brochus Harold and Winterbottom, 1999
Gobiodon ceramensis (Bleeker, 1852)
Gobiodon citrinus (Rüppell, 1838)
Gobiodon fulvus Herre, 1927
Gobiodon heterospilos Bleeker, 1856
Gobiodon histrio (Valenciennes, 1837)
Gobiodon micropus Günther, 1861
Gobiodon okinawae Sawada, Arai and Abe, 1972
Gobiodon quinquestrigatus (Valenciennes, 1837)
Gobiodon reticulatus Playfair, 1866
Gobiodon rivulatus (Rüppell, 1830)
Gobiodon spilophthalmus Fowler, 1944
Gobiodon unicolor (Castelnau, 1873)
Gobiopsis angustifrons Lachner and McKinney, 1978
Gobiopsis aporia Lachner and McKinney, 1978
Gobiopsis bravoi (Herre, 1940)
Gobiopsis exigua Lachner and McKinney, 1979
Gobiopsis macrostoma Steindachner, 1861
Gobiopsis malekulae (Herre, 1935)
Gobiopsis quinquecincta (Smith, 1931)
Gobiopsis springeri Lachner and McKinney, 1979
Gobiopsis woodsi Lachner and McKinney, 1978
Hazeus otakii Jordan and Snyder, 1901
Hetereleotris poecila (Fowler, 1946)
Isthmogobius baliurus (Valenciennes, 1837)
Istigobius decoratus (Herre, 1927)
Istigobius diadema (Steindachner, 1877)
Istigobius goldmanni (Bleeker, 1852)
Istigobius hoesei Murdy and McEachran, 1982
Istigobius nigroocellatus (Günther, 1873)
Istigobius ornatus (Rüppell, 1830)
Istigobius rigilius (Herre, 1953)
Istigobius spence (Smith, 1947)
Kellogella quindecimfasciata (Fowler, 1946)
Kelloggella cardinalis Jordan and Seale, 1906
Lobulogobius morrigu Larson, 1983
Lobulogobius omanensis Koumans, 1944
Lophogobius bleekeri Popta, 1921
Lotilia graciliosa Klausewitz, 1960
Lubricogobius ornatus Fourmanoir, 1966
Lubricogobius pumilis Larson and Hoese, 1980
Luposicya lupus Smith, 1959
Macrodontogobius wilburi Herre, 1936
Mahidolia mystacina (Valenciennes, 1837)
Mangarinus waterousi Herre, 1943
Myersina crocatus Wongratana, 1975
Myersina lachneri Hoese and Lubbock, 1982
Myersina macrostoma Herre, 1934
Myersina nigrivirgata Akihito and Meguro, 1983
Myersina papuensis (Peters, 1876)
Obliquogobius cometes (Alcock, 1890)
Oplopomops diacanthus (Schultz, 1943)
Oplopomus caninoides (Bleeker, 1852)
Oplopomus oplopomus (Valenciennes, 1837)
Opua nephodes Jordan, 1925
Palutrus pruinosa (Jordan and Seale, 1906)
Palutrus scapulopunctatus (Beaufort, 1912)
Parachaeturichthys ocellatus (Day, 1873)
Parachaeturichthys polynema (Bleeker, 1853)
Paragobiodon echinocephalus (Rüppell, 1830)
Paragobiodon lacunicolus (Kendall and Goldsborough, 1911)
Paragobiodon melanosomus (Bleeker, 1852)
Paragobiodon modestus (Regan, 1908)
Paragobiodon xanthosomus (Bleeker, 1852)
Parkraemeria ornata Whitley, 1951
Phyllogobius platycephalops (Smith, 1964)
Platygobiopsis akihito Springer and Randall, 1992
Pleurosicya annandalei Hornell and Fowler, 1922
Pleurosicya australis Larson, 1990
Pleurosicya bilobata (Koumans, 1941)
Pleurosicya boldinghi Weber, 1913
Pleurosicya carolinensis Larson, 1990
Pleurosicya coerulea Larson, 1990
Pleurosicya elongata Larson, 1990
Pleurosicya fringilla Larson, 1990
Pleurosicya labiata (Weber, 1913)
Pleurosicya micheli Fourmanoir, 1971
Pleurosicya mossambica Smith, 1959
Pleurosicya muscarum (Jordan and Seale, 1906)
Pleurosicya plicata Larson, 1990
Pleurosicya prognatha Goren, 1984
Pleurosicya spongicola Larson, 1990
Priolepis agrena Winterbottom and Burridge, l993
Priolepis ailina Winterbottom and Burridge, l993
Priolepis aithiops Winterbottom and Burridge, l992
Priolepis aureoviridis (Gosline, 1959)
Priolepis cincta (Regan, 1908)
Priolepis compita Winterbottom, 1985
Priolepis fallacincta Winterbottom and Burridge, l992
Priolepis inhaca (Smith, 1949)
Priolepis kappa Winterbottom and Burridge, 1991
Priolepis nocturna (Smith, 1957)
Priolepis nuchifasciata (Günther, 1973)
Priolepis pallidicincta Winterbottom and Burridge, l993
Priolepis profunda (Weber, 1909)
Priolepis semidoliata (Valenciennes, 1837)
Priolepis squamogena Winterbottom and Burridge, 1989
Priolepis sticta Winterbottom and Burridge, l992
Priolepis triops Winterbottom and Burridge, l993
Psammogobius biocellatus (Valenciennes, 1837)
Psilogobius mainlandi Baldwin, 1972
Psilogobius prolatus Watson and Lachner, 1985
Signigobius biocellatus Hoese and Allen, 1977
Silhouettea evanida Larson and Miller, 1986
Silhouettea hoesei Larson and Miller, 1986
Silhouettea insinuans Smith, 1959
Silhouettea nuchipunctatus (Herre, 1934)
Stonogobiops nematodes Hoese and Randall, 1982
Stonogobiops xanthorhinica Hoese and Randall, 1982
Sueviota aprica Winterbottom and Hoese, 1988
Sueveiota lachneri Winterbottom and Hoese, 1988
Sueviota larsonae Winterbottom and Hoese, 1988
Tomiyamichthys latruncularia (Klausewitz, 1974)
Tomiyamichthys oni (Tomiyama, 1936)
Trimma benjamini Winterbottom, 1996
Trimma caesiura Jordan and Seale, 1906
Trimma caudimaculata Yoshino and Araga, 1977
Trimma emeryi Winterbottom, 1985
Trimma grammistes (Tomiyama, 1936)
Trimma griffithsi Winterbottom, 1984
Trimma hoesei Winterbottom, 1984
Trimma macrophthalma (Tomiyama, 1936)
Trimma mendelssohni (Goren, 1978)
Trimma naudei Smith, 1956
Trimma necopinna Whitley, 1959
Trimma okinawae (Aoyagi, 1949)
Trimma rubromaculata Allen and Munday, 1995
Trimma sheppardi Winterbottom, 1984
Trimma striata (Herre, 1945)
Trimma taylori Lobel, 1979
Trimma tevegae Cohen and Davis, 1969
Trimma unisquamis (Gosline, 1959)
Trimmatom eviotops (Schultz, 1943)
Trimmatom macropodus Winterbottom, 1989
Trimmatom nanus Winterbottom and Emery, 1981
Trimmatom sagma Winterbottom, 1989
Trimmatom zapotes Winterbottom, 1989
Valenciennea alleni Hoese and Larson, 1994
Valenciennea bella Hoese and Larson, 1994
Valenciennea decora Hoese and Larson, 1994
Valenciennea helsdingeni (Bleeker, 1858)
Valenciennea immaculata Ni, 1981
Valenciennea limicola Hoese and Larson, 1994
Valenciennea longipinnis (Lay and Bennett 1839)
Valenciennea muralis (Valenciennes, 1837)
Valenciennea parva Hoese and Larson, 1994
Valenciennea puellaris (Tomiyama, 1955)
Valenciennea randalli Hoese and Larson, 1994
Valenciennea sexguttata (Valenciennes, 1837)
Valenciennea strigata (Broussonet, 1782)
Valenciennea wardi (Playfair, 1866)
Vanderhorstia ambanoro (Fourmanoir, 1957)
Vanderhorstia flavilineata Allen and Munday, 1995
Vanderhorstia lanceolata Yanagisawa, 1978
Vanderhorstia mertensii Klausewitz, 1974
Vanderhorstia ornatissima Smith, 1959
*Awaous acritosus Watson, 1994
*Awaous guamensis Valenciennes, 1837
*Awaous litturatus (Steindachner, 1860)
*Awaous melanocephalus (Bleeker, 1849)
*Awaous ocellaris (Broussonet, 1782)
Brachygobius doriae (Günther, 1868)
Brachygobius kabiliensis Inger, 1958
Brachygobius xanthozona (Bleeker, 1849)
Calamiana kabilia (Herre, 1940)
Calamiana mindora (Herre, 1945)
Calamiana variegata (Peters, 1869)
Chlamydogobius ranunculus (Larson, 1995)
Eugnathogobius microps Smith, 1931
Gnatholepis anjerensis (Bleeker, 1851)
Gnatholepis cauerensis (Bleeker, 1853)
Gnatholepis deltoides (Seale, 1901)
Gnatholepis inconsequens Whitley, 1958
Gnatholepis scapulostigma Herre, 1953
Gobiopterus brachypterus (Bleeker, 1855)
Gobiopterus chuno (Hamilton, 1822)
Gobiopterus luzonensis (Smith, 1902)
Gobiopterus panayensis (Herre, 1944)
Gobiopterus semivestitus (Munro, 1949)
Hemigobius hoevenii (Bleeker, 1851)
Hemigobius mingi (Herre, 1936)
Mugilogobius cavifrons (Weber, 1909)
Mugilogobius chulae (Smith, 1932)
Mugilogobius fusca (Herre, 1940)
Mugilogobius fusculus (Nichols, 1951)
Mugilogobius mertoni (Weber, 1911)
Mugilogobius notospilus (Gunther, 1877)
Mugilogobius platynotus (Gunther, 1861)
Mugilogobius platystomus (Gunther, 1872)
Mugilogobius rambaiae (Smith, 1945)
Mugilogobius stigmaticus (De Vis, 1884)
Oligolepis acutipennis (Valenciennes, 1837)
Oligolepis jaarmani (Weber, 1913)
Oligolepis stomias (Smith, 1941)
Oxyurichthys auchenolepis Bleeker, 1867
Oxyurichthys cornutus McCulloch and Waite, 1918
Oxyurichthys lonchotus (Jenkins, 1903)
Oxyurichthys microlepis (Bleeker, 1849)
Oxyurichthys notonema (Weber, 1909)
Oxyurichthys ophthalmonema (Bleeker, 1856)
Oxyurichthys papuensis (Valenciennes, 1837)
Oxyurichthys takagi Pezold, 1998
Oxyurichthys tentacularis (Valenciennes, 1837)
Oxyurichthys uronema (Weber, 1909)
Pandaka pusilla Herre, 1927
Pandaka pygmaea Herre, 1927
Pandaka rouxi (Weber, 1911)
Pandaka trimaculata Akihito and Meguro, 1975
Pandaka lidwilli (McCulloch, 1917)
Pseudogobius avicennia (Herre, 1940)
Pseudogobius javanicus (Bleeker, 1856)
Pseudogobius melanosticta (Day, 1876)
Pseudogobius poicilosoma (Bleeker, 1849)
Redigobius balteatus (Herre, 1935)
Redigobius bikolanus (Herre, 1927)
Redigobius chrysosomus (Bleeker, 1875)
Redigobius macrostomus (Gunther, 1861)
Rediogbius roemeri (Weber, 1911)
*Rhinogobius giurinus (Rutter, 1897)
*Stenogobius alleni Watson, 1991
*Stenogobius beauforti (Weber, 1908)
*Stenogobius blokzeyli (Bleeker, 1861)
*Stenogobius caudimaculosus Watson, 1991
*Stenogobius fehlmanni Watson, 1991
*Stenogobius gymnopomus (Bleeker, 1853)
*Stenogobius hoesei Watson, 1991
*Stenogobius ingeri Watson, 1991
*Stenogobius kyphosus Watson, 1991
*Stenogobius lachneri Watson, 1994
*Stenogobius
laterisquamatus (Weber, 1905)
*Stenogobius marinus Watson, 1991
*Stenogobius marqueti Watson, 1991
*Stenogobius ophthalmoporus (Bleeker, 1853)
*Stenogobius psilosinionus Watson, 1991
*Stenogobius randalli Watson, 1991
*Stenogobius squamosus Watson, 1991
*Stenogobius zurstrasseni (Popta, 1912)
Stigmatogobius borneensis (Bleeker, 1851)
Stigmatogobius pleurostigma (Bleeker, 1849)
Stigmatogobius sadanundio (Hamilton, 1822)
Stigmatogobius sella (Steindachner, 1881)
Subfamily OXUDERCINAE
Apocryptodon madurensis (Bleeker, 1849)
Boleophthalmus birdsongi Murdy, 1989
Boleophthalmus boddarti (Pallas, 1770)
Boleophthalmus caeruleomaculatus McCulloch and Waite, 1918
Oxuderces dentatus (Eydoux and Souleyet, 1848)
Oxuderces wirzi (Koumans, 1937)
Parapocryptes serperaster (Richardson, 1846)
Periophthalmodon freycineti (Valenciennes, 1824)
Periophthalmodon schlosseri (Pallas, 1770)
Periophthalmodon septemradiatus (Hamilton, 1822)
Periophthalmus argentilineatus Valenciennes, 1837
Periophthalmus chrysospilos Bleeker, 1852
Periophthalmus gracilis Eggert, 1935
Periophthalmus kalolo Lesson, 1830
Periophthalmus malaccensis Eggert, 1935
Periophthalmus minutus Eggert, 1935
Periophthalmus novaeguineaensis Eggert, 1935
Periophthalmus novemradiatus (Hamilton, 1822)
Periophthalmus weberi Eggert, 1935
Pseudapocryptes borneensis (Bleeker, 1855)
Pseudapocryptes lanceolatus (Bloch and Schneider, 1801)
Scartelaos histophorus (Valenciennes, 1837)
Subfamily SICYDIINAE
*Lentipes crittersius Watson and Allen, 1999
*Lentipes dimetrodon Watson and Allen, 1999
*Lentipes watsoni Allen, 1997
*Lentipes whittenorum Watson and Kottelat, 1994
*Sicyopterus cynocephalus (Valenciennes, 1837)
*Sicyopterus eudentatus Parenti and Maciolek, 1993
*Sicyopterus hageni Popta, 1921
*Sicyopterus japonicus (Tanaka, 1909)
*Sicyopterus lividis Parenti and Maciolek, 1993
*Sicyopterus longifilis De Beaufort, 1912
*Sicyopterus
macrostetholepis (Bleeker, 1853)
*Sicyopterus marquesensis Fowler, 1932
*Sicyopterus microcephalus (Bleeker, 1854)
*Sicyopterus micrurus (Bleeker, 1853)
*Sicyopterus ouwensi Weber, 1913
*Sicyopterus parvei (Bleeker, 1853)
*Sicyopterus pugnans (Ogilvie-Grant, 1884)
*Sicyopterus taeniurus (Gunther, 1877)
*Sicyopterus wichmanni De Beaufort, 1912
*Sicyopus auxilimentus Watson and Kottelat, 1994
*Sicyopus bitaeniatus Maugé et al., 1986
*Sicyopus discordipinnis Watson, 1995
*Sicyopus fehlmanni Parenti and Maciolek, 1993
*Sicyopus leprurus Sakai and Nakamura, 1979
*Sicyopus multisquamatus De Beaufort, 1912
*Sicyopus mystax Watson and Allen, 1999
*Sicyopus nigriradiatus Parenti and Maciolek, 1993
*Sicyopus zosterophorum (Bleeker, 1856)
*Stiphodon allen Watson, 1996
*Stiphodon astilbos Ryan, 1986
*Stiphodon atratus Watson, 1996
*Stiphodon atropurpureus (Herre, 1927)
*Stiphodon birdsong Watson, 1996
*Stiphodon caeruleus Parenti and Maciolek, 1993
*Stiphodon elegans (Steindachner, 1859)
*Stiphodon hydroreibatus Watson, 1999
*Stiphodon larson Watson, 1996
*Stiphodon olivaceus Watson and Kottelat, 1995
*Stiphodon ornatus Meinken, 1974
*Stiphodon pelewensis Herre, 1936
*Stiphodon rutilaureus Watson, 1996
*Stiphodon semoni Weber, 1895
*Stiphodon stevensoni (Jordan and Seale, 1906)
*Stiphodon surrufus Watson and Kottelat, 1995
*Stiphodon zebrinus Watson, Allen and Kottelat, 1998
